Ecologist
Ecosystem Health and Restoration, Public Lands Ecology Section, Lake Michigan Ecological Research Station

Education:
B.S. (Biology), Stephen F. Austin State University, 1971
M.S. (Limnology), Stephen F. Austin State University, 1973
Ph.D. (Wildlife and Fishery Sciences), Texas A&M University, 1979

Research Interests:
E. coli, eutrophication, contaminants, environmental assessment, biomonitoring indicators

Mailing Address:
USGS Great Lakes Science Center
Lake Michigan Ecological Research Station
1100 North Mineral Springs Road
Porter, IN 46304

E-mail: rwhitman@usgs.gov

Phone: (219) 926-8336, ext. 424

Fax: (219) 929-5792

Byappanahalli, Muruleedhara N., Tao Yan, Matthew J. Hamilton, Satoshi Ishii, Roger S. Fujioka, Richard L. Whitman, Michael J. Sadowsky. 2012. The population structure of Escherichia coli isolated from subtropical and temperate soils. Science of the Total Environment 417/418: 273-279.
Contribution # 1677 [Abstract]

Ge, Zhongfu, Richard L. Whitman, Meredith B. Nevers, Mantha S. Phanikumar, and Muruleedhara N. Byappanahalli. 2012. Nearshore hydrodynamics as loading and forcing factors for Escherichia coli contamination at an embayed beach. Limnology and Oceanography 57(1): 362-381.
Contribution # 1664 [Abstract] [PDF 3.49 MB]

Ge, Zhongfu., Richard L. Whitman, Meredith B. Nevers, and Mantha S. Phanikumar. 2012. Wave-induced mass transport affects daily Escherichia coli fluctuations in nearshore water. Environmental Science & Technology 46(4): 2204-2211.
Contribution # 1683 [Abstract]

Badgley, Brian D., John Ferguson, Amy Vanden Heuvel, Gregory T. Kleinheinz, Colleen M. McDermott, Todd R. Sandrin, Julie Kinzelman, Emily A. Junion, Muruleedhara N. Byappanahalli, Richard L. Whitman, and Michael J. Sadowsky. 2011. Multi-scale temporal and spatial variation in genotypic composition of Cladophora-borne Escherichia coli populations in Lake Michigan. Water Research 45(2): 721-731.
Contribution # 1607 [Abstract]

Nevers, Meredith B. and Richard L. Whitman. 2011. Efficacy of monitoring and empirical predictive modeling at improving public health protection at Chicago beaches. Water Research 45(4): 1659-1668.
Contribution # 1627 [Abstract]

Nevers, Meredith B. and Richard L. Whitman. 2011. Beach monitoring criteria: Reading the fine print. Environmental Science and Technology 45(24): 10315–10321.
Contribution # 1670 [Abstract]

Byappanahalli, Muruleedhara N., Richard L. Whitman, Dawn A. Shively, and Meredith B. Nevers. 2010. Linking non-culturable (qPCR) and culturable enterococci densities with hydrometeorological conditions. Science of the Total Environment 408(16): 3096-3101.
Contribution # 1586 [Abstract]

Dorevitch, Samuel, Nicholas J. Ashbolt, Ferguson M. Christobel, Roger Fujioka, Charles D. McGee, Jeffrey A. Soller, and Richard L. Whitman. 2010. Meeting report: knowledge and gaps in developing microbial criteria for inland recreational waters. Environmental Health Perspectives 118(6): 871-876.
Contribution # 1583 [Abstract]

Ge, Zhongfu, Meredith B. Nevers, David J. Schwab, and Richard L. Whitman. 2010. Coastal loading and transport of Escherichia coli at an embayed beach in Lake Michigan. Environmental Science & Technology 44(17): 6731-6737.
Contribution # 1601 [Abstract]

Nevers, Meredith B., and Richard L. Whitman. 2010. Policies and practices of beach monitoring in the Great Lakes, USA: a critical review. Journal of Environmental Monitoring 12(3): 581-590.
Contribution # 1574 [Abstract]

Thupaki, Pramod, Mantha S. Phanikumar, Dmitry Beletsky, David J. Schwab, Meredith B. Nevers, and Richard L. Whitman. 2010. Budget analysis of Escherichia coli at a southern Lake Michigan beach . Environmental Science & Technology 44(3): 1010-1016.
Contribution # 1562 [Abstract]

Verhougstraete, Marc P., Muruleedhara N. Byappanahalli , Joan B. Rose, and Richard L. Whitman. 2010. Cladophora in the Great Lakes: impacts on beach water quality and human health. Water Science and Technology 62(1): 68-76.
Contribution # 1578 [Abstract]

Whitman, Richard L., Meredith B. Nevers, Katarzyna Przybyla-Kelly, and Muruleedhara N. Byappanahalli. 2010. Physical and biological factors influencing environmental sources of fecal indicator bacteria in surface water. Pages 111-134 in Sadowsky, Michael J. and Richard L. Whitman, eds. The Fecal Bacteria. ASM Press, Washington, DC.
Contribution # 1599 [Abstract]

Whitman, Richard L., Zhongfu Ge, Meredith B. Nevers, Alexandria B. Boehm, Eunice C. Chern, Richard A. Haugland, Ashley M. Lukasik, Marirosa Molina, Kasia Przybyla-Kelly, Dawn A. Shively, Emily M. White, Richard G. Zepp, and Muruleedhara N. Byappanahalli. 2010. Relationship and variation of qPCR and culturable enterococci estimates in ambient surface waters are predictable. Environmental Science & Technology 44(13): 5049-5054.
Contribution # 1591 [Abstract]

Byappanahalli, Muruleedhara N. and Richard L. Whitman. 2009. Clostridium botulinum type E occurs and grows in the alga Cladophora glomerata. Canadian Journal of Fisheries and Aquatic Sciences 66(6): 879-882.
Contribution # 1531 [Abstract]

Byappanahalli, Muruleedhara N., Richard Sawdey, Satoshi Ishii, Dawn A. Shively, John A. Ferguson, Richard L. Whitman, and Michael J. Sadowsky. 2009. Seasonal stability of Cladophora-associated Salmonella in Lake Michigan watersheds. Water Research 43(3): 806-814.
Contribution # 1514 [Abstract]

Nevers, Meredith B., Dawn A. Shively, Gregory T. Kleinheinz, Colleen M. McDermott, William Schuster , Vinni Chomeau, and Richard L. Whitman. 2009. Geographic relatedness and predictability of Escherichia coli along a peninsular beach complex of Lake Michigan. Journal of Environmental Quality 38(6): 2357-2364.
Contribution # 1538 [Abstract]

Whitman, Richard L., Katarzyna Przybyla-Kelly, Dawn A. Shively, Meredith B. Nevers, and Muruleedhara N. Byappanahalli. 2009. Hand-mouth transfer and potential for exposure to E. coli and F+ coliphage in beach sand, Chicago, Illinois. Journal of Water and Health 7(4): 623-629.
Contribution # 1534 [Abstract]